
Topics
What is bovine somatotropin (bST)?
How did scientists develop bST?
How does bST affect milk production?
What are the benefits and risks of bST?
How is bST regulated?
What are the controversies concerning the use
of bST?
What lies ahead?
For more information
[Topics]
What is bovine somatotropin (bST)?
Bovine somatotropin is a growth hormone found in cattle. The word "bovine"
refers to cattle, and the word "somatotropin" refers to the name
of the hormone. Hormones are chemicals that are secreted by glands within
the body. They are natural substances that affect the way the body operates.
Bovine somatotropin, abbreviated as bST, is a protein hormone produced in
cattle by the pituitary gland located at the base of the animal's brain.
A hormone similar to bST is produced in all species of animals. This hormone
is important for growth, development, and other bodily functions of all
animals. In the 1930s, it was discovered that injecting bST into lactating
(milk-producing) cows significantly increased milk production.
[Topics]
How did scientists develop bST?
Until recently, the only source of bST was from the pituitary glands of
slaughtered cattle. There were only small quantities of bST available, and
it was very expensive.
Now, the new science of biotechnology makes it possible to work with DNA,
the part of a cell that contains the genetic information for an animal or
a plant. Scientists have determined which gene in cattle controls or "codes"
for the production of bST. They have removed this gene from cattle and inserted
it into a bacterium called Escherichia coli. This bacterium, which
is found in the intestinal tract of humans and animals, acts like a tiny
factory and produces large amounts of bST in controlled laboratory conditions.
The bST produced by the bacteria is purified and then injected into cattle
(figure 1).
The movement of a gene from one organism to another, in this case from
the pituitary gland of a cow to a bacterial organism, is called "recombinant
DNA technology." Several Food and Drug Administration (FDA) approved
drugs, including insulin for the treatment of diabetes and tissue plasminogen
activator (TPA) for the treatment of heart attacks in people, are produced
in a similar way.
[Topics]
How does bST affect milk production?
To affect a cow's milk production, bST must be injected into the animal
on a regular basis, similar to the way insulin must be regularly injected
into people who have certain types of diabetes. Feeding bST to cows will
not work. Amino acids and peptides are the building blocks of proteins.
The hormone bST is a complex protein that is immediately broken down into
small, inactive amino acids and peptides and rendered ineffective when it
enters a cow's digestive system. How often a cow must be injected with bST
will depend on whether a bST product can be developed that releases the
hormone gradually over a long period of time.
Milk yields are significantly increased when cows are injected with bST,
although not as much as some reports in popular newspapers and magazines
suggest. The exact details of how bST increases milk production are not
known, but it is thought that blood flow to the cow's mammary (milk-producing)
gland is increased. The blood carries an increased amount of nutrients available
for milk production. More nutrients are extracted from the blood by the
mammary gland, which improves efficiency of milk production. Feed efficiency
(pounds of milk produced per pound of feed consumed) is improved because
more milk is produced and the proportion of feed used for body maintenance
is decreased. The actual amount of feed consumed by bST-treated cows increases,
helping the cow meet the increased nutrient demands.
Milk production in bST-treated cows increases from 4.8 to 11.2 pounds per
day. Feed efficiency improves from 2.7 to 9.3 percent (Peel, et al.). Table
1 summarizes the results of 32-week treatments of cows injected with
bST in several states and foreign countries.
Misinformation provided by some groups gives the impression that there
is controversy about the biology of somatotropin. However, 800 reports on
20,000 treated cows have yielded remarkably consistent results worldwide
(Bauman).
Researchers have summarized several bST trials and found a milk production
increase of 8.4 pounds per day (Bauman). They estimated that, depending
on how the dairy operation is managed, average increased milk production
is expected to range from 8.5 to 17.6 percent.
It is difficult to predict how individual cows will respond to bST. A higher
response is seen when treatment is started after the cow has been producing
milk for 101 days, rather than when treatment is started on days 57-100
after calving. The response of cows treated in early lactation is less (Bauman).
Cows that have had more than one calf show a greater increase in milk production
than do first lactation heifers (Peel, et al.). Milk yield gradually increases
for the first few days after bST treatment begins. A maximum increase is
seen in about six days. To meet the needs for this increased milk production,
treated cows consume from 10 to 20 percent more grain and forage.
Normally, cows reach their peak milk production 7-9 weeks after lactation
begins. Milk quantity then slowly declines throughout the remainder of lactation.
The ability of cows to maintain relatively high levels of milk production
throughout lactation is called "persistency." The major response
of cows treated with bST is a significant improvement in persistency. The
normal decrease in milk yield as lactation progresses is markedly reduced.
Quality of management, including health programs, milking practices, nutrition,
cow condition, and environmental conditions will be major factors in the
response to bST.
[Topics]
What are the benefits and risks of bST?
The commercial use of bST in dairy cattle is controversial and has stirred
heated debate among the dairy industry, activist groups, and consumers.
Effects on cow health
The physiological effects of bST treatment are the same as those seen in
any high-producing cow. Nutrition, health programs, environment, and milking
technique must be appropriate for the use of bST or results will be disappointing.
On many farms, the management changes instituted by producers as they are
preparing to use bST will probably cause a greater increase in milk production,
efficiency, and profitability than actual use of bST. In the initial stages
of use, producers will be encouraged to use bST on cows that have been in
lactation for at least 100 days, are in good physical condition, pregnant,
and are free from health problems such as mastitis or infertility.
Concern has been expressed regarding the effect of bST on reproduction.
The optimum calving interval of 12-13 months may lengthen because bST can
extend the time that cows efficiently produce milk. Dairy Herd Improvement
Association (DHIA) records show that higher milk-producing herds have lower
conception rates than lower producing herds (Ferguson and Skidmore). This
negative effect on reproduction is seen in cows treated with bST and is
associated with increased milk production. However, some people believe
that a longer calving interval could benefit the health of bST- treated
cows, since many health problems of dairy cows are associated with calving
and rebreeding. The ability of a cow to reproduce is affected by her physical
condition, nutrition, health, and level of milk production.
Few research studies have investigated the physiological effects of bST
on the functioning of the ovaries and pituitary gland. Cows receiving dosages
of bST far beyond what will be used in practice have shown an adverse effect
on estrous activity (the time when an animal is capable of being bred).
This effect is not seen when cows receive low to average dosages of bST.
High dosages of bST are reported to increase the death rate of calf embryos,
so starting a cow on bST during early pregnancy should probably be avoided
(Ferguson and Skidmore). This effect is not seen at recommended dosages.
The effect of bST on reproduction will have to be monitored closely in individual
herds.
Several research studies have shown that bST is not associated with increased
mammary infections (mastitis) (Ferguson and Skidmore). Other studies have
shown an increase in mammary gland infections when bST is used, but the
increase is what would be expected with increased production. The length
of a cow's gestation (pregnancy), calf birth weight, calf survival rate,
and calf growth are not influenced by using the product. Some early reports
indicated an increased incidence of twins, but later reports failed to confirm
this.
Milk composition and safety
Consumer advocates and others have expressed concern about the safety of
milk from bST-treated cows. All milk contains natural bST that is produced
by the cow. Milk from bST-treated cows also contains the same amounts of
injected bST and no differences can be measured compared to untreated cows.
There are four forms of natural bST, and each has a chain of either 190
or 191 amino acids. The recombinant bST that is injected into cows has 191
amino acids. The biological activity of commercial bST is identical to naturally
produced bST.
Studies indicate that both natural bST produced by the cow and bST produced
by recombinant DNA techniques are immediately broken down into inactive
amino acids and peptides in the digestive tract when they are consumed by
humans. In contrast, steroid hormones such as estrogens, progesterones,
and anabolic steroids are smaller, ring-like structures that are absorbed
from the digestive tract and are biologically active in humans. This is
not the case with bST in milk, whether it is produced naturally by the cow
or by recombinant DNA technology (Barbano and Lynch).
Studies show that bovine somatotropin is inactive in humans. During the
1950s, natural bST produced by cows was injected into children with growth
defects in an attempt to encourage growth. There was no effect, probably
because the bovine somatotropin protein molecule differs from human somatotropin
(human growth hormone) by about 30 percent of the amino acid sequences.
Milk composition from bST-treated cows has been thoroughly investigated
(Barbano and Lynch). The characteristics of milk from bST-treated cows are
within the normal range of variation of milk from untreated cows. During
the first 28 days of treatment, milk fat increases and milk protein decreases
slightly. After longer treatment, cows adjust their nutrient intake and
the normal balance is reestablished. An increase in nonprotein nitrogen
and whey protein and a decrease in casein have been observed after long-term
bST administration. This difference is not always significant, and the effect
on cheese yield would probably be minor, if any. One study showed a slight
increase in unsaturated compared to saturated fat. The difference was small,
but suggested a healthier product from bST treatment. No differences in
free fatty acids have been observed. Cholesterol levels are in the range
of normal milk composition. Insulin-like growth factor I increases by up
to two-fold in milk from treated cows, but it is still well within the range
for both bovine and human milk. No differences in flavor have been found.
The National Institute of Health has concluded that milk from bST-treated
cows is essentially the same as from untreated cows, and there is no difference
in safety of the products.
Economics
The potential economic effect of bST on the family dairy farm has generated
heated debate. The Animal Health Institute, an organization of drug and
vaccine manufacturers, maintains that the use of bST will be of equal value
to any size farm (Milligan). They contend that use of the product will favor
the good dairy manager, regardless of farm size.
Estimates of the effect of bST on dairy production have probably been exaggerated.
The United States Department of Agriculture estimates that the use of bST
could lead to a 2 to 5 percent increase in national milk production within
five years, or about the increase seen yearly without the use of bST. This
increase would be in addition to the normal milk production increase per
cow.
In most dairy herds, bST will not be used in cows until they have been in
lactation for about 100 days. It will not be used in cows with chronic health
or fertility problems. It is expected that bST will be used less in heifers
than in adult cows. If 50 percent of farmers adopt the use of bST, and it
is used in 60 percent of the lactation days per user herd, milk production
will increase about 3.5 percent (assuming an average per cow production
increase of 15 percent). Many well-managed dairy herds increase per cow
production more than this on an annual basis by using improved management
and genetics. For most herds, a farmer who requests a thorough herd analysis
by a competent nutritionist and veterinarian and then follows their recommendations
will achieve a larger increase in milk production than by using bST alone.
Failure to adopt proven technology is a problem throughout the dairy industry.
Almost 60 percent of cows are bred by mating to a bull, rather than by artificial
insemination from proven sires with superior genetic performance. Only 50
percent of U.S. dairy producers use DHIA management information and records
to improve production.
The government milk price support system tends to make prediction of the
effect of bST on milk prices difficult. It is true, however, that efficient
managers in areas of the country with higher milk prices benefit more from
application of technology and increased production.
It is argued that large commercial dairy operations can begin using new
technologies such as bST more easily, rapidly, and efficiently than smaller
operations. Sophisticated record keeping and division of labor may make
timely injection of cows with bST more feasible for these larger operations.
Other demands on the time and management skills of typical Midwest dairy
producers who have diversified farming operations may make new technology
more difficult to implement.
Others argue that smaller producers with direct owner control of the herd
can manage individual cows better and will see a greater production increase
from the use of bST. It is not automatically true that larger herds are
better managed and, therefore, will benefit more from bST.
There is no question that consumer loss of confidence in the quality of
milk produced by using bST, whether the reasons are logical or not, would
reduce milk consumption and have a negative economic impact on the dairy
industry. This is a major concern of dairy producers. Relative to milk quality,
bST appears to be neutral. It neither improves nor harms quality. Consumers
would gain with bST technology since milk production costs may decline due
to improved efficiency. The ultimate effect of use of bST on consumption
is unknown.
[Topics]
How is bST regulated?
The United States Food and Drug Administration (FDA) is responsible for
regulating the use of bST since it is an animal drug and because milk and
meat are food products. Several commercial companies have submitted data
to the FDA, asking for approval to use bST in dairy cows to increase milk
production.
Before approval of bST for use in dairy herds, the FDA allowed the consumption
of milk and meat from animals that received bST as part of the experimental
testing process. Such approval is often granted during the process of license
approval of animal products. Milk from treated cows has been judged safe
because bST is biologically inactive in humans and is a protein hormone
that is digested and destroyed by gastric enzymes when it is consumed. Each
company seeking approval for bST has to demonstrate that bST has zero biological
activity in milk when it is consumed. "The FDA has found no pertinent
information indicating that food derived from bST-treated cows is unsafe."
(Review). It is not required that producers withhold milk from the market
for a certain period of time after test herd cows have been treated with
bST.
Federal law prohibits the social and economic effects of a product from
affecting the FDA's decision whether to approve its use or not. The FDA
must determine if a product is safe, pure, potent, and effective. Producers
can decide whether a product is economical or useful. Approval by the FDA
does not mean that a product must be used, but only that it can be used,
if desired.
Testing required by the FDA
Before any new product can be approved, companies must demonstrate its effectiveness
under actual use conditions in several geographic locations. Fifty cows
per herd are required for bST approval. Three dosages of bST were used for
the studies submitted to the FDA. The quality control of bST used in the
test herds was monitored and all procedures to be used were approved by
the FDA before the testing began. The majority of the tests in the approval
process were performed by independent scientists at university laboratories
and farms or in commercial herds.
Cows were injected with bST at various times during the lactation period.
The effectiveness of the drug and its safety for the first and later lactation
periods were monitored. Milk yield was calculated on a 3.5 percent fat basis.
Milk composition, including fat, crude protein, lactose, calcium, and phosphorus,
were measured about once per week. Daily feed intake was measured in the
test herds. Body condition and health were monitored throughout the studies.
The effect of bST treatment on reproduction was evaluated, including breeding
cycles, conception rates, number of breedings per conception, length of
time from calving to the next conception, abortions, incidence of twins,
calving difficulties, and stillbirths. The weight, growth, and health of
calves during the first four weeks of life were monitored. Monthly somatic
cell counts, as a measure of mastitis, were required. The sites where bST
was injected were monitored for any signs of adverse reactions.
To evaluate safety, companies had to use one, three, and five times the
expected dosage level of bST for two consecutive lactations in one of their
test herds. Heifers born to treated cows were raised through breeding age
and monitored for abnormalities. Companies seeking approval for bST were
also required to prove that its use was not harmful to the environment.
First bST product approval granted by FDA
On November 5, 1993, the FDA announced approval of a bST product, the animal
drug sometribove, for increasing milk production in dairy cows. The Monsanto
Company of St. Louis, Missouri, developed the drug. However, the drug could
not be used immediately due to a 90-day moratorium imposed by Congress during
the summer of 1993. The moratorium was designed to give the White House
Office of Management and Budget time to study possible consumer reaction
and the drug's impact on the dairy industry.
The FDA approval also carried with it some provisions to deal with antibiotic
residue concerns. In September 1992, the General Accounting Office reported
that the FDA had found evidence in submitted clinical trials that bST-treated
cows have a slightly increased incidence of mastitis. This report raised
concerns that antibiotic treatments for mastitis could lead to increased
antibiotic residues in milk. States require milk to be tested for drug residues.
Milk found to have unsafe levels of residues must be discarded.
Although an FDA advisory committee concluded in March 1993 that adequate
safeguards exist to prevent unsafe levels of antibiotic residues from entering
the milk supply, additional steps were taken to ensure that any unsafe residues
in the milk of bST-treated cows are detected before the milk or its products
are marketed.
According to a news release issued by the U.S. Department of Health and
Human Services (HHS News), Monsanto agreed to a post-approval monitoring
program that includes:
- A two-year tracking system of milk production and drug residues in 21
top dairy states that will periodically compare the amount of milk discarded
after bST is marketed to the amount discarded prior to approval.
- A 12-month comparison of the proportion of milk discarded due to positive
drug tests between bST-treated and untreated herds.
- A reporting system to monitor all bST use and follow up on all complaints.
- The use of sometribove in 24 commercial dairy herds will be specifically
monitored for mastitis, animal drug use, and the resulting loss of milk.
The FDA has concluded that it has no legal basis to require special labeling
of food products derived from bST-treated cows. Food companies may voluntarily
label their products, provided the information is truthful and not misleading
to consumers.
[Topics]
What are the controversies concerning the use of bST?
The most intense controversy surrounding approval of bST for use in dairy
cows has occurred in major dairy producing states in the Great Lakes and
New England areas. Representatives of the dairy industry are concerned about
the ultimate economic effect on producers.
Consumer and environmental advocacy groups have expressed opposition based
on concerns about milk quality and the use of biotechnology in general.
Dairy producers
Some producers are afraid that they will not be able to keep up with new
technologies and they will suffer economically as a result. Others feel
that a product such as bST will work to the disadvantage of producers in
the Great Lakes States and the Northeast. Natural resistance to new technology
adoption and a fear of genetic engineering techniques cause some producers
to resist the approval and use of bST.
Special interest groups
Activist groups with a variety of agendas and motives have addressed the
bST issue. Some have stated that milk from treated cows may not be safe
after all, and more testing is needed. Others see this as a scare tactic
to delay or block the use of bST and undermine consumer confidence in milk
from bST-treated cows. Some animal rights groups see the use of animals
for food, under any circumstances, as inhumane or a violation of those animals'
"rights." Others have stated that cows have a right not to be
injected with bST.
Others
Other opposing arguments state that the FDA does no independent testing
of its own, but only monitors the studies of the companies seeking approval.
The persistent oversupply of milk and dairy products has also been cited
as a reason to block the use of bST. Some dairy farmers oppose the use of
bST but feel they would have no choice but to use the product in their own
herd in order to stay competitive if bST came into general use (McDermott).
Table 2 summarizes arguments for and against
the use of bST in the categories of food safety and its effect on the number
and size of farms.
[Topics]
What lies ahead?
Use of bST will have a significant effect on the research and development
investment in agricultural biotechnology by commercial firms. Universities
will be expected to provide unbiased scientific information.
Patience, tolerance, and understanding will be required by educators, extension
workers, and other professionals in agriculture who work with groups that
either support or oppose implementation of technology such as bST.
[Topics]
For more information
Dairy News and Information Center, 2233 Wisconsin Avenue, N.W., Suite 500,
Washington, D. C. 20007. Tel. toll free 1-800-343-2479.
Nolan R. Hartwig, D.V.M. Extension Veterinarian, 2270E Veterinary Medicine
Complex, Iowa State University, Ames, Iowa 50011. Tel. (515) 294-8790.
Agricultural Biotechnology: Issues and Choices. Copyright 1991 by Purdue
Research Foundation. Available from Agricultural Communication Service,
Media Distribution Center, Purdue University, West Lafayette, Indiana 47907.
References
Barbano, David M. and Joanna M. Lynch. "Milk from bST Treated Cows:
Composition and Manufacturing Properties." Advanced Technologies Facing
the Dairy Industry: bST. Cornell University. Nov. 10-11, 1989.
Bauman, Dale E. "Biology of Bovine Somatotropin." Advanced Technologies
Facing the Dairy Industry: bST. Cornell University. Nov. 10-11, 1989. p.
1-8.
Ferguson, James D., and Andrew Skidmore. "Bovine Somatotropin - Reproduction
and Health." Advanced Technologies Facing the Dairy Industry: bST.
Cornell University. Nov. 10-11, 1989. p. 57-66.
HHS News. U.S. Department of Health and Human Services. Nov. 5, 1993.
McDermott, Keyron. Editorial, The Des Moines Register, Des Moines, Iowa,
May 3, 1990.
Milligan, Robert A. "Profitability of bST Technology." Advanced
Technologies Facing the Dairy Industry: bST. Cornell University. Nov. 10-11,
1989. p. 36-48.
Peel, C. J.; D. L. Hurd; K. S. Madsen; and G. de Kerchove. Monsanto Agricultural
Company. In Proceedings, Monsanto Technical Symposium, Oct. 24, 1989. The
Monsanto Company, St. Louis, Missouri.
Review of Bovine Somatotropin. FDA Veterinarian. March/April 1990, Vol.
V, No. II.
Stanton, B. F., and A. M. Novalovic. "The Impact of bST Technologies:
The Pros and Cons." Advanced Technologies Facing the Dairy Industry:
bST. Cornell University. Nov. 10-11, 1989. p. 142-148.
Written by Nolan R. Hartwig, D.V.M. Iowa State University Extension Veterinarian,
and Glenda D. Webber, Office of Biotechnology, Iowa State University.
North Central Regional Extension Publications are subject to peer review
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March, 1994

North Central Regional
Extension Publication
NCR #488